During this section, CO2 is free of malate and re-fixed by Rubisco. For this to happen, malic acid is re-mobilized by passive efflux and decarboxylated. There are two various pathways for malate decarboxylation, relying on the species. One begins with the release of CO2 by the conversion of malate to pyruvate by NAD-ME, followed by the regeneration of PEP by PPDK (Kondo et al., 2000).

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In a second pathway, NAD-MDH converts malate to OAA, followed by PEPCK, regenerating PEP, and releasing CO2 . Finally, phase IV takes place late within the afternoon once malate reserves have been depleted. At this time, stomata open once more and permit for the uptake of CO2, which is processed immediately by Rubisco (Dodd et al., 2002).

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